Boddole Zer is the supreme commander of the 4,795,122-ship Boddole Zer Main Fleet belonging to the main antagonists, the Zentradi. Golg Boddole Zer ordered his subordinate Britai Kridanik to track down and recover the Supervision Army spaceship which had crashlanded on Earth, thus starting Space War I. After the failures of Britai and his other subordinate Moruk Lap Lamiz to capture the SDF-1 Macross and their subsequent "cultural contamination" by the miclones, Bodole Zer ordered all 4,795,122 ships in the Boddole Zer Main Fleet to fold to Earth and destroy the SDF-1 Macross, the defecting Zentradi forces, and the Earth itself in 2010. Boddole Zer was killed when his massive flagship was destroyed in an unorthodox tactic by the SDF-1 Macross, but not before he ordered the obliteration of the Earth's surface with less than one million human survivors in the Battle with the Boddole Zer Main Fleet.